Why has my rabbits started peeing on the bed

My rabbits leo and Charlie have been letterbox trained for around 5 months over the past month they have gained access to the bed, they have been very good with it, no poo or pee. However over the past week leo had decided to start peeing on the bed and Charlie followed.

There has been no changed to their letterbox location or the litter I use, the only change in their routine is they go outside a little earlier as it has gotten hot.
When I check the camera to see who peed on the bed 4/5 times it is leo.
I saw that he only peed on the bed where Charlie was sat, he came up to him bit him and then peed there, leo is very stubborn and has a habit of peeing on thigs after getting his nails trimmed or doing something he doesn't like ( cleaning). I'm unsure as to why Charlie peed on the bed as he is the well behaved one.
I bought a new duvet and covers to hopefully get rid of any pee smell and it hasn't worked,
Is leo just being difficult or is it something I have done to cause this?

What can I do to get him to stop peeing on the bed, they live in my room with me and the only other option is for them to live outside if they can't be in my room.
 
Are they neutered? Rabbits usually mark territory more if unneutered. If they are both unneutered males you are likely to have a big problem on your hands at some stage and this behaviour may be the start of it (I am assuming they are both male from the names). Male rabbits will literally fight to the death once they reach sexual maturity if not neutered, even if they are getting on well now. If they are neutered then a vet trip may be needed to figure out if there's a physical cause.
 
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