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Rabbits - face pulling

I was playing with theo today, giving him nose rubs, and he seemed to be enjoying it, but was pulling subtle little faces

Its hard to describe, but he was slighly pulling one cheek in, if he were a human, I would describe it as the face you would make if you were sucking your molar teeth on one side of your mouth

He wasn't distressed at all, and he does this occasionally

Just wondering what it means
 
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